Source: Warhammer Fantasy: 7th Edition

Different Sized Bases within Units
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It is quite common for a Lizardmen army to include units containing models with different sized bases such as a Slann in a Temple Guard unit or one or more Kroxigor in a Skink unit - so it is worth some clarification as to how this affects ranks and rank bonuses.

The best way to handle rank bonuses is to count the model on the larger base as the same number of models that would normally occupy that space. So, a Slann in a Temple Guard unit would count as four models two in each rank - when determining rank bonus (see example diagram below).

Furthermore, by the normal Warhammer rules, a rear rank cannot be larger than the one in front. Following the above principle, if there were two Kroxigors in a unit of Skinks, and only three Skinks were left in the front rank, one of the Kroxigor would have to move forward to the front rank.

combined-units-figure-1
Fig 1: This unit has a rank bonus of +2. The first rank consists of six models, the second and third ranks count as having six models because the Slann occupies the space of two in each rank. The fourth rank only has four models, too few to count towards the rank bonus.
combined-units-figure-2
Fig 2: This mixed unit of Skinks and Kroxigors starts out with a rank bonus of +2. Even though it has lost a Skink from its third rank, this rank and the second still count as having at least five models in them because the Kroxigors occupy the space of two models in each. Later on in the battle, only six Skinks remain. These form the front rank, while the Kroxigor rank up behind them. However, if three more Skinks are killed, one of the Kroxigors has to step forward, because a unit's rear rank may not have more models than any other rank.

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